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- Aplysia spp. in the Ria Formosa lagoon (S. Portugal): are they important herbivores?Publication . Sprung, Martin; Aníbal, J.; Santos, Rui; Vieira, V.The impact of the opistobranchs Aplysia fasciata and Aplysia depilans on green algae in the Ria Formosa lagoon has been assessed by a combination of field observations and laboratory experiments.