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- Off the shelf: regulation of fish blood pH through ionocyte membrane remodelling and protein translocation without additional synthesiPublication . Guerreiro da Costa Guerreiro, Pedro Miguel; Canario, AdelinoOcean acidification (OA), together with warming and deoxygenation, forms the “deadly trio,” a combination that reduces ecosystems’ resilience, makes marine biodiversity vulnerable, and is considered a main contributor to global mass extinctions in the Earth’s history (1).