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- The role of lithic technology in shaping mobility and decision-making: the case of Ararat-1 CavePublication . Cabrita Nora, David André; Frahm, Ellery; Oikonomou, Ioannis A.K.; Karampaglidis, Theodoros; Gasparyan, Boris; Petrosyan, Artur; Buller, Ariel MalinskyStudies of lithic technological organization have progressed from static typological classifications to analyses of dynamic processes of tool production, use, and discard. These analyses reveal the intricate interplay of human behavior and environmental adaptation. This paper investigates lithic technology, emphasizing the dichotomy of curated and expedient technologies by examining the differential strategies employed in raw material acquisition, tool production, and discard. It also explores how environmental, economic, and mobility factors interplay and shape lithic assemblages. Focusing on Ararat-1 cave, a late Middle Paleolithic site in the Armenian Highlands, we conducted an integrative techno-typological study of 1770 lithic artefacts spanning five stratigraphic units, identifying distinct raw material reduction strategies for obsidian and chert. Obsidian artefacts exhibited a focus on retouching and rejuvenation, indicative of curated technologies associated with high residential mobility, while the chert showed evidence of primary reduction and blank production, reflecting expedient strategies. By integrating module flake analysis and reviewing the Whole Assemblage Behavioral Indicators (WABI) framework, suggesting the addition of smaller debitage, we highlighted dual mobility patterns reflected in the site assemblage. This approach offers a refined understanding of lithic technological systems, providing insights into MIS 3 mobility and site function in the Armenian Highlands.
- The non-adult burials of cabeço da amoreira, muge (Portugal): recent discoveries and new insights into mesolithic funerary practicesPublication . Coutinho-Nogueira, Dany; Godinho, Ricardo Miguel; Gaspar, Rosa Ramos; Andre, Lino; Bicho, Nuno; Cascalheira, João; Gonçalves, Célia; Umbelino, CláudiaCabeço da Amoreira, a key Mesolithic shell mound in the Muge region (central Portugal) provides critical insights into the funerary and socio-cultural practices of Western Europe’s last hunter-gatherer-fisher populations. Recent systematic excavations have provided new data into biological diversity and funerary practices, particularly in relation to non-adult individuals. This study focuses on three non-adult burials uncovered since 2019. By employing state-of-the-art imaging techniques, we digitally recorded, virtually excavated, and analysed the biological profiles of these individuals. Our results demonstrate a notable variability in funerary practices, including differences in burial orientation, the presence or absence of artefacts, and even the construction of a purposeful burial feature for one individual. These discoveries contribute significantly to the growing body of research on Mesolithic funerary traditions, illustrating the complexity of social and ritual behaviours in southwestern Iberia’s last hunter-gatherer-fisher societies and opening new avenues for comparative research on childhood and burial practices in prehistory.
- Evolution of craniofacial shape in relation to sexual dimorphism in theropithecus and papioPublication . Ossorio, Ángeles; Figueroa-Torrejón, Ambra; Buston, Rodrigo; Caldon, Matteo; Mathe, Jacinto; Doria, Giuliano; Gippoliti, Spartaco; Volta, Antonella; Silva, María Joana Ferreira da; Bobe, René; Carvalho, Susana; Capelli, Cristian; Martínez, Felipe I.Introduction: Sexual dimorphism in cranial morphology is a significant aspect of primate evolution, providing insights into evolutionary pressures and mating systems in different species. This study focuses on cranial sexual dimorphism in Papio and Theropithecus, two closely related genera within the tribe Papionini.Methods: Using geometric morphometric techniques, we analyzed 570 cranial specimens from both genera, with data sourced from various studies and repositories. Thirty craniofacial landmarks were defined and analyzed through Geometric Morphometrics tools to evaluate shape variation.Results: Our findings reveal distinct morphological clusters for each genus and sex, with Papio and Theropithecus exhibiting significant sexual dimorphism. The results distinguish genera and sex-based groups, indicating differential impacts of size on shape across groups. The findings suggest that while sexual dimorphism is stable in magnitude within each genus, the specific morphological manifestations differ.Discussion: This research advances our understanding of the evolutionary mechanisms driving sexual dimorphism and emphasizes the need for further studies to explore the genetic and environmental factors influencing these differences. The innovative approach and comprehensive dataset provide a robust framework for future investigations into primate cranial morphology and its evolutionary implications.
- Intensive resource exploitation in Late Neolithic Iberia: bone marrow and subsistence changes at Branqueiras, central-coastal PortugalPublication . Costa, Cláudia; Fernandes Martins, Maria João; Cardoso, Vera; Cardoso, Guilherme; Carvalho, António Manuel Faustino deThe exploitation of bone marrow has long been recognised as a vital component of past human subsistence strategies, particularly for its nutritional value. While this practice is well studied in hunter-gatherer societies and in communities facing harsh environmental conditions, it is less understood in prehistoric farming societies and temperate environments. This paper presents the results of bone marrow exploitation at the Late Neolithic short term site of Branqueiras, located in the temperate central-coastal region of Portugal. The faunal assemblage reveals a significant percentage of intentional long bone breakage for marrow extraction, a practice that appears to have been overlooked in earlier and later periods. The site’s short-term occupation, radiocarbon-dated to 2900–2700 cal BC, coincides with a hiatus at the nearby long-term settlement of Leceia and occurred during a period of climate change characterised by reduced rainfall, hot summers, and cold winters. This study provides key insights into human adaptation to climate stress within established farming communities, a topic often underexplored in Neolithic–Chalcolithic contexts. The authors propose that the Branqueiras community repre sents a shift in human behaviour, as itinerant groups adapted to arid conditions by intensively exploiting animal resources, including domestic caprines, swine, wild game, and molluscs, while agricultural practices and cattle herding declined. In this context, bone marrow became a crucial source of nutrients, playing a significant role in subsistence strategies during this challenging period Phenolic compounds.
- Material matters: raw material influences stone tool performance in capuchin monkeysPublication . O’Malley, Theo D. R.; Slania, Nora E.; Falótico, Tiago; Torre, Ignacio de la; Luncz, Lydia V.; Reeves, Jonathan S.; Proffitt, TomosIdentifying the conditions that facilitate and shape tool use is a central focus in the field of human evolution and animal behaviour. Particular interest lies in the use of stone hammers by nonhuman primates to open encased food sources. It is widely theorized that similar behaviours were used by early hominins and provided a foundation for the emergence of stone knapping. Environmental factors are thought to be important in shaping the emergence and progression of tool use. However, there is limited information on whether access to different types of raw tool material for hammerstones and anvils affects the reliability or efficiency with which tool users exploit encased resources. Here, we experimentally provide wild capuchins, Sapajus libidinosus, in Brazil with raw materials differing in hardness. Materials were sourced globally from primate and hominin tool use sites. We measured the reliability and efficiency with which monkeys could crack nuts when using different raw materials, and how these metrics changed over the course of the experiment. We further reported variations in the durability of different raw materials, which directly relates to how long a tool remains useable. Our results showed that differences in capuchin nut-cracking performance were largely driven by the ability of the tool material to stabilize the nut on the anvil. Furthermore, there was wide variation in anvil durability during use. These differences appeared to be driven by multiple tool characteristics, including hardness, surface texture and anvil and hammerstone mass. When compared with similar studies, our results also suggest that stone properties, particularly hardness, may have differing effects on nut-cracking outcomes across species. Overall, the differences in raw material performance and durability seen here, respectively, highlight how local raw materials may influence the selective costs and benefits of tool use behaviours, and the accumulation of tools within the landscape.
- Predictors of baboon sleep site selection in gorongosa national parkPublication . Lewis‐Bevan, Lynn; Hammond, Philippa; Carvalho, Susana; Biro, DoraObjectives: This research aimed to understand how sleep site selection compared to other study sites in baboons living in a lowpredator density, highly seasonal environment. We compared baboon troops in two distinct habitat types with different seasonal influences within the park, one that flooded annually and one that did not. We compared their sleep site use, reuse, and location relative to home range boundaries and areas of interest (AOIs) with each other and baboons in other areas to understand whether season, habitat familiarity, or position in the home range influenced sleep site choice. Methods: Using GPS collar data taken at 15-min intervals from four gray-footed chacma baboons (Papio ursinus griseipes) in Gorongosa National Park, Mozambique, we established the location of sleep sites, home range boundaries, and AOIs, or places where the baboons repeatedly stopped for more than 15 min. Study subjects ranged either in dense woodland or in a seasonally flooded alluvial floodplain. We used a linear mixed-effects model to predict sleep site reuse based on distance to the habitat edge and AOIs, and Wilcoxon signed-rank tests to determine if morning or evening AOIs influenced sleep site location. We counted the number of reuses of each sleep site before and after the flooding period and compared this data to data in other baboon study sites. Results: We found that, as in other study sites with less seasonality and higher predation risk, baboons in Gorongosa change sleep site frequently and utilize multiple sleep sites throughout their home range, although they more often use sleep sites closer to the center of their home ranges. However, unlike other studies, we found that the location of the last AOI of the day more strongly predicted sleep site location than the first AOI of the next day in one troop, with baboons traveling further from their sleep site to their first AOI in the morning than from their last evening AOI to the sleep site. Conclusions: Despite high seasonality and low predator density, baboons in Gorongosa National Park changed sleep sites frequently, as do other studied baboon troops in areas with high nocturnal predation rates. In addition, their propensity to sleep closer to the last AOI of the day may imply that they plan their daily paths toward their chosen sleep site, or that they sleep opportunistically at the end of the day. This study provides a baseline of behavioral data for comparison to other sites and future work in Gorongosa, where predator density continues to rise since the time of the study.
- Hominin glacial-stage occupation 712,000 to 424,000 years ago at Fordwich Pit, Old Park (Canterbury, UK)Publication . Key, Alastair; Clark, James; Lauer, Tobias; Bates, Jennifer; Sier, Mark-Jan; Nichols, Claire; Martín-Ramos, Carmen; Cebeiro, Adela; Williams, Eleanor; Kim, Sunghui; Stileman, Finn; Mika, Anna; Pope, Matthew; Bridgland, David; Redhouse, David; Leonardi, Michela; Smith, Geoff M.; Proffitt, TomosFew high-latitude archaeological contexts are older than marine isotope stage (MIS) 15 and even fewer provide evidence of early human occupation during a glacial period. New discoveries at Old Park, Canterbury (UK), provide evidence of both the oldest accessible artefact-bearing sediment in northern Europe and cold-stage adaptation. Radiometric and palaeomagnetic dating places the earliest suggested occupation of this site between 773 thousand years ago (ka) and 607 ka, with hominin presence inferred during MIS 17–16. Two additional artefact-bearing stratigraphic units, dated to around 542 ka and 437 ka, strongly align with the MIS 14 and 12 cold stages, respectively. The latter unit contains convincing evidence of glacial-stage occupation by Acheulean hominins; fresh, unabraded flakes (including biface-thinning) between clearly defined glacial-aged sediments displaying mixed grassland palaeoenvironmental evidence. An historically collected assemblage of more than 330 handaxes is argued to be derived from both the MIS 17–16 and MIS 12 sediments, providing evidence of the earliest known Acheulean bifaces in northern Europe, and re-occupation by Acheulean populations 200,000 years later. Together, Old Park provides evidence for Lower Palaeolithic hominins reoccupying a location over several mid-Pleistocene MIS cycles, early human presence above 51° latitude during a glacial stage and handaxe production in northern Europe from MIS 17 to 16.
- Assessing neanderthal occupation duration: faunal evidence from sub-unit IIIb of teixoneres cave (Barcelona, Spain)Publication . Lubrano, Valentina; Rufà, Anna; Blasco, Ruth; Rivals, Florent; Rosell, JordiDetermining the duration of occupations at Middle Palaeolithic archaeological sites is essential to enhancing our understanding of Neanderthal behaviour and subsistence strategies. Identifying occupation length in these contexts presents significant complexity and is often ambiguous due to the palimpsest nature of the archaeological sites. This study examines faunal specimens from sub-Unit IIIb of Teixoneres Cave, focusing on both external and internal site areas, to estimate the duration of human occupations. Using zooarchaeological and taphonomic parameters, we evaluate the effectiveness of these methods for inferring occupation length. Our analysis shows that faunal remains—such as evidence of hunting, butchering, and carcass transport—are essential for reconstructing Neanderthal subsistence strategies and for understanding whether the site was used during one short-term occupation or across several distinct episodes over time. While certain results diverge from the features currently used in literature to define occupation duration, the spatial distribution of lithic artefacts, from sub-Unit IIIb at Teixoneres Cave support the hypothesis of short, repeated human occupations. This analysis focuses on applying and critically examining some of those theoretical assumptions, particularly those related to taphonomic and archaeozoological criteria. In this context, Teixoneres Cave serves as the dataset through which we test whether a real archaeological case supports the theoretical framework proposed in Lubrano et al. (2025). These findings contribute to ongoing discussions on the complexities of interpreting archaeological palimpsests and the methodological challenges involved in estimating the duration of Neanderthal occupations. Furthermore, the data place sub-Unit IIIb of Teixoneres within the broader framework of Middle Palaeolithic cave sites characterized by short-term human occupations in settings frequently utilized by large carnivores.
- The genetic history of Portugal over the past 5,000 years.Publication . Roca-Rada, Xavier; Davidson, Roberta; Williams, Matthew P.; Villalba-Mouco, Vanessa; Carvalho, António Manuel Faustino de; Ravishankar, Shyamsundar; Collen, Evelyn; Haarkötter, Christian; Taufik, Leonard; Cuesta-Aguirre, Daniel R.; Tente, Catarina; Calleja, Álvaro M Monge; MacRoberts, Rebecca Anne; Melo, Linda; Purnomo, Gludhug A.; Souilmi, Yassine; Tobler, Raymond; Cunha, Eugénia; Tereso, Sofia; Matos, Vítor M. J.; Fernandes, Teresa Matos; Maurer, Anne-France; Silva, Ana Maria; Carvalho, Pedro C.; Llamas, Bastien; Teixeira, João C.Recent ancient DNA studies uncovering large-scale demographic events in Iberia have presented very limited data for Portugal, a country located at the westernmost edge of continental Eurasia. Here, we present the most comprehensive collection of Portuguese ancient genome-wide data, from 67 individuals spanning 5000 years of human history, from the Neolithic to the nineteenth century. We identify early admixture between local hunter-gatherers and Anatolian-related farmers in Neolithic Portugal, with a northeastern-southwestern gradient of increasing Magdalenian-associated ancestry persistence in Iberia. This profile continues into the Chalcolithic, though Bell Beaker-associated sites reveal Portugal's first evidence of Steppe-related ancestry. Such ancestry has a broader demographic impact during the Bronze Age, despite continuity of local Chalcolithic genetic ancestry and limited Mediterranean connections. The village of Idanha-a-Velha emerges in the Roman period as a site of significant migration and interaction, presenting a notably diverse genetic profile that includes North African and Eastern Mediterranean ancestries. The Early Medieval period is marked by the arrival of Central European genetic diversity, likely linked to migrations of Germanic tribes, adding to coeval local, African, and Mediterranean influences. The Islamic and Christian Conquest periods show strong genetic continuity in northern Portugal and significant additional African admixture in the south. The latter remains stable during the post-Islamic period, suggesting enduring African influences. We reveal dynamic patterns of migration in line with cultural exchange across millennia, but also the persistence of local ancestries. Our findings integrate genetic information with historical and archeological data, enhancing our understanding of Iberia's biological and cultural heritage.
- Open science, communication, and collaboration for the future of palaeoproteomicsPublication . Dekker, Joannes Adrianus Antonius; Peters, Carli; Winter, Rachel M.; Collins, Matthew J.; Dickinson, Marc R.; Harvey, Virginia L.; Hill, Eden; Nair, Bharath; Tsutaya, Takumi; Viñas-Caron, Laura C.; Warinner, Christina; Welker, Frido; Fagernäs, ZandraPalaeoproteomics focuses on studying the past by analysing proteins preserved in historical, archaeological, and palaeontological materials. Over the past fifteen years, the number of publications and research questions in this field has grown substantially, as has the number of researchers and research groups utilising palaeoproteomic methods. PAASTA (Palaeoproteomics And Archaeology, Society for Techniques and Advances) is an international early career researcher-driven community, which aims to bring together researchers from all subdisciplines of palaeoproteomics by providing a platform for open communication. Here, we discuss ways in which this fast-growing research field can benefit from adopting principles of open science and increased communication and collaboration between researchers. We approach this question from all stages of a typical research project, from experimental design and sample selection, to data analysis and raw data sharing. Although we acknowledge that each of these guidelines and recommendations may not be applicable to every research project, we aim to provide a foundation which, when applied collectively, will strengthen the future of palaeoproteomics.